Early Triassic

The climate during the Early Triassic Epoch (especially in the interior of the supercontinent Pangaea) was generally arid, rainless and dry and deserts were widespread; however the poles possessed a temperate climate. The pole-to-equator temperature gradient was temporally flat during the Early Triassic and may have allowed tropical species to extend their distribution poleward. This is evidenced by the global distribution of ammonoids.^[13] The extremely hot ocean temperatures facilitated extremely powerful hurricanes that frequently hit the coast of North China.^[14]

The mostly hot climate of the Early Triassic may have been caused by late volcanic eruptions of the Siberian Traps,^[15][8] which had probably triggered the Permian-Triassic extinction event and accelerated the rate of global warming into the Triassic.^[16] Studies suggest that Early Triassic climate was very volatile, punctuated by a number of relatively rapid global temperature changes, marine anoxic events, and carbon cycle disturbances,^[17][18][19] which led to subsequent extinction events in the aftermath of the Permian-Triassic extinction event.^[20][21][22] On the other hand, an alternative hypothesis proposes these Early Triassic climatic perturbations and biotic upheavals that inhibited the recovery of life following the P-T mass extinction to have been linked to forcing driven by changes in the Earth's obliquity defined by a roughly 32.8 thousand year periodicity with strong 1.2 million year modulations. According to proponents of this hypothesis, radiometric dating indicates that major activity from the Siberian Traps ended very shortly after the end-Permian extinction and did not span the entire Early Triassic epoch, thus not being the primary culprit for the climatic changes throughout this epoch.^[23]

Fauna and flora

The Triassic Period opened in the aftermath of the Permian–Triassic extinction event. The massive extinctions that ended the Permian Period (and with that the Paleozoic Era) caused extreme hardships for the surviving species.

The Early Triassic Epoch saw the biotic recovery of life after the biggest mass extinction event of the past, which took millions of years due to the severity of the event and the harsh Early Triassic climate.^[24] Many types of corals, brachiopods, molluscs, echinoderms, and other invertebrates had disappeared. The Permian vegetation, which was dominated by Glossopteris in the Southern Hemisphere, ceased to exist.^[25] Other groups, such as Actinopterygii, appear to have been less affected by this extinction event^[26] and body size was not a selective factor during the extinction event.^[27][28] Animals that were most successful in the Early Triassic were those with high metabolisms.^[29] Different patterns of recovery are evident on land and in the sea. Early Triassic faunas lacked biodiversity and were relatively homogeneous due to the effects of the extinction. The ecological recovery on land took 30 million years, well into the Late Triassic.^[30] Two Early Triassic lagerstätten stand out due to their exceptionally high biodiversity, the Dienerian aged Guiyang biota^[31] and the earliest Spathian aged Paris biota.^[32]

Terrestrial biota

The most common land vertebrate was the small herbivorous synapsid Lystrosaurus. Often interpreted as a disaster taxon (although this view was questioned^[33]), Lystrosaurus had a wide range across Pangea. In the southern part of the supercontinent, it co-occurred with the non-mammalian cynodonts Galesaurus and Thrinaxodon, early relatives of mammals. First archosauriforms appeared, such as Erythrosuchus (Olenekian-Ladinian).^[34] This group includes the ancestors of crocodiles and dinosaurs (including birds). Fossilized foot prints of dinosauromorphs are known from the Olenekian.^[35] The Early Triassic entomofauna is very poorly understood because of the paucity of insect fossils from this epoch.^[36]

The flora was gymnosperm-dominated at the onset of the Triassic, but changed rapidly and became lycopod-dominated (e.g. Pleuromeia) during the Griesbachian-Dienerian ecological crisis. This change coincided with the extinction of the Permian Glossopteris flora.^[25] In the Spathian subage, the flora changed back to gymnosperm and pteridophyte dominated.^[37] These shifts reflect global changes in precipitation and temperature.^[25][20] Floral diversity was overall very low during the Early Triassic, as plant life had yet to fully recover from the Permian-Triassic extinction.^[38]

Microbially induced sedimentary structures (MISS) are common in the fossil record of North China in the immediate aftermath of the Permian-Triassic extinction, indicating that microbial mats dominated local terrestrial ecosystems following the Permian-Triassic boundary. The regional prevalence of MISS is attributable to a decrease in bioturbation and grazing pressure as a result of aridification and temperature increase.^[39] MISS have also been reported from Early Triassic fossil deposits in Arctic Canada.^[40] The disappearance of MISS later in the Early Triassic has been interpreted as a signal of increased bioturbation and recovery of terrestrial ecosystems.^[39]

Aquatic biota

In the oceans, the most common Early Triassic hard-shelled marine invertebrates were bivalves, gastropods, ammonoids, echinoids, and a few articulate brachiopods. Conodonts experienced a revival in diversity following a nadir during the Permian.^[41] The first oysters (Liostrea) appeared in the Early Triassic. They grew on the shells of living ammonoids as epizoans.^[42] Microbial reefs were common, possibly due to lack of competition with metazoan reef builders as a result of the extinction.^[43] However, transient metazoan reefs reoccurred during the Olenekian wherever permitted by environmental conditions.^[44] Ammonoids show blooms followed by extinctions during the Early Triassic.^[45]

Aquatic vertebrates diversified after the extinction:

• Fishes: Many species of fish had a worldwide distribution during the Early Triassic.^[27] Typical Triassic ray-finned fishes, such as Australosomus, Birgeria, Bobasatrania, Boreosomus, Pteronisculus, Parasemionotidae and Saurichthys appeared close to the Permian-Triassic boundary, whereas neopterygians (including stem teleosts) diversified later during the Triassic, though the pattern of the Triassic diversification of bony fishes is not well understood due to a taphonomic megabias (Spathian-Bithynian Gap, SBG) in the late Early Triassic and early Middle Triassic.^[46] The earliest large durophagous neopterygian is known from this gap,^[47] suggesting an early onset in feeding specializations of this group. Most bony fish reached large body sizes during the Early Triassic.^[27] Coelacanths show a peak in their diversity during this epoch,^[48] including new modes of life, such as the fork-tailed Rebellatrix.^[49] Chondrichthyes are represented by Hybodontiformes like Palaeobates, Omanoselache, Lissodus, some Neoselachii, as well as a few last survivors of the Eugeneodontida (Caseodus, Fadenia).^[50]

• Amphibians: Relatively large, marine temnospondyl amphibians, such as Aphaneramma or Wantzosaurus, were geographically widespread during the Induan and Olenekian ages. The fossils of these crocodile-shaped amphibians were found in Greenland, Spitsbergen, Pakistan and Madagascar.^{[citation needed]}

• Reptiles: In the oceans, first marine reptiles appeared during the Early Triassic.^[51] Their descendants ruled the oceans during the Mesozoic. Hupehsuchia, Ichthyopterygia and sauropterygians are among the first marine reptiles to enter the scene in the Olenekian (e.g. Cartorhynchus, Chaohusaurus, Utatsusaurus, Hupehsuchus, Grippia, Omphalosaurus, Corosaurus). Other marine reptiles such as Tanystropheus, Helveticosaurus, Atopodentatus, placodonts or the thalattosaurs followed later in the Middle Triassic.^[51] The Anisian aged ichthyosaur Thalattoarchon was one of the first marine macropredators capable of eating prey that was similar in size to itself, an ecological role that can be compared to that of modern orcas.^[52]

Fossil gallery

• 
  Early Triassic brittle stars (echinoderms)
• 
  Fossils of the bivalve Claraia clarai
• 
  Early Triassic ammonoid Hedenstroemia
• 
  Fossil of the Early Triassic neopterygian Candelarialepis argentus
• 
  Early Triassic Hupehsuchus fossil in the Paleozoological Museum of China
• 
  Skull of the Early Triassic archosauriform Erythrosuchus
• 
  Lystrosaurus hedini skeleton at the Museum of Paleontology in Tübingen

• Geologic time scale
• Triassic
• Mass extinction

• Martinetto, Edoardo; Tschopp, Emanuel; Gastaldo, Robert, eds. (2020). Nature through Time: Virtual field trips through the Nature of the past. Springer International Publishing. ISBN 978-3-030-35057-4.

• GeoWhen Database – Early Triassic
• Palaeos (archived 2 January 2006)
• scotese