龍蝨科
鞘翅目的一科昆虫
（重定向自Dytiscidae）
龍蝨科（学名：Dytiscidae），亦作龍虱科，為水甲蟲的一種，隸屬於鞘翅目之下的肉食亞目。生活在田野、水溝、小溪等水體中，是一種掠食性水生昆蟲。其學名源於希臘文δυτικός（dytikos），意思就是「肉食性的潛水甲蟲」。本科物種幾乎見於全世界所有淡水棲境，但亦有部分於落葉堆中棲息[1]。成蟲體長一般在1和2.5 cm（0.4—1.0英寸）之間，儘管物種間的差異會比較大。當中見於歐洲的Dytiscus latissimus及巴西的Megadytes ducalis有最大的體型，可分別長至4.5 cm（1.8英寸）到4.75 cm（1.9英寸）[1][2]。與之相反，擁有最小體型的物種應當為棲息於澳大利亞地下水中的Limbodessus atypicali，只有約0.9 mm（0.035英寸）長[1]。大多數物種的顏色都在深棕色、黑色或深橄榄绿，某些亚科的物種會在其邊沿帶有金黃色的條狀斑紋。
龍蝨科 化石时期：侏儸紀晚期至今 | |
|---|---|
_-_(imago),_Berkenwoude,_the_Netherlands.jpg) | |
| Cybister lateralimarginalis | |
科学分类  | |
| 界： | 动物界 Animalia |
| 门： | 节肢动物门 Arthropoda |
| 纲： | 昆虫纲 Insecta |
| 目： | 鞘翅目 Coleoptera |
| 亚目： | 肉食亞目 Adephaga |
| 总科： | 龍蝨總科 Dytiscoidea |
| 科： | 龍蝨科 Dytiscidae Leach, 1815 |
| 模式属 | |
| 龙虱属 Dytiscus Linnaeus，1758 | |
| 亞科、族與属 | |
| 多样性 | |
| 至少160個屬，約4000物種 | |
食用
本科物種的成蟲，特别是Cybister屬的物種，均為可食用昆蟲。在美國内华达州洪堡乾湖的一個山洞内，還在當地史前人類遺下的粪化石内發現有Cybister explanatus被食用的殘骸[3]。在墨西哥，他們在吃C. explanatus時會以墨西哥夹饼夾着來吃。在日本多個地區（例如：長野縣）有以日本龍蝨（Cybister japonicus）為食的習慣；在中华人民共和国广东省，除了日本龍蝨以外，亦會食用C. bengalensis、C. guerini、C. limbatus、C. sugillatus、C. tripunctatus等物種。D. marginalis很可能也包括在內，但這個物種的肉很少。也有報告指在台灣、泰国及新幾內亞亦有食用本科物種的習慣[4]。
大但在陸地上很慢，成年後也不是特別兇猛，它們也被許多中型鳥類、哺乳動物和其他大型捕食者津津有味地吃掉。幼蟲通常比較安全，由於它們的保護色和通過水柱逃跑的能力；它們可能相當難抓，可能成為小池塘的頂級掠食者。
系統分類
- Acilius Leach, 1817
- Aethionectes Sharp, 1882
- Africodytes Biström, 1988
- Africophilus Guignot, 1948
- Agabetes Crotch, 1873
- Agabinus Crotch, 1873
- 豆龍蝨屬 Agabus Leach, 1817[6]，端毛龙虱属
- Agametrus Sharp, 1881
- Agaporomorphus Zimmermann, 1921
- Aglymbus Sharp, 1880
- Agnoshydrus Biström, Nilsson & Wewalka, 1997
- Allodessus Guignot, 1953
- 彩斑龍蝨屬 Allopachria Zimmermann, 1924
- Amarodytes Régimbart, 1900
- Amurodytes Fery & Petrov, 2013
- Andex Sharp, 1882
- Andonectes Guéorguiev, 1971
- Anginopachria Wewalka, Balke & Hendrich, 2001
- Anodocheilus Babington, 1841
- Antiporus Sharp, 1882
- Australphilus Watts, 1978
- Austrelatus Shaverdo, Hájek, Hendrich, Surbakti, Panjaitan & Balke, 2023
- Austrodytes Watts, 1978
- Barrethydrus
- Barretthydrus Lea, 1927
- Barretthydrus geminatus Lea, 1927
- Barretthydrus stepheni Watts, 1978
- Barretthydrus tibialis Lea, 1927
- Batrachomatus Clark, 1863
- Belladessus K.B.Miller & Short, 2015
- Bidessini Sharp, 1880
- Bidessodes Régimbart, 1895
- Bidessonotus Régimbart, 1895
- Bidessus Sharp, 1882
- Boreonectes (Fery & Ribera, 2018) Fery & Ribera, 2018
- Boreonectes Angus, 2010
- Borneodessus Balke, Hendrich, Mazzoldi & Biström, 2002
- Brachyvatus Zimmermann, 1919
- Brancuporus Hendrich, Toussaint & Balke, 2014
- Bunites Spangler, 1972
- Calicovatellus K.B.Miller & Lubkin, 2001
- Canthyporus Zimmermann, 1919
- Capelatus Turner & Bilton, 2015
- Caperhantus Balke, Hájek & Hendrich, 2017
- Carabdytes Balke, Hendrich & Wewalka, 1992
- Carabhudrus
- Carabhydrus Watts, 1978
- Carrhydrus
- Celina Aubé, 1837
- Chostonectes Sharp, 1880
- Clarkhydrus Fery & Ribera, 2018
- Clemnius Villastrigo, Ribera, Manuel, Millán & Fery, 2017
- Clypeodytes Régimbart, 1894
- Coelambus Thomson, 1860
- Coelambus cribrarius (Scudder, 1900)
- Coelambus derelictus (Scudder, 1900)
- Coelambus disjectus (Scudder, 1900)
- Coelambus dzieduszyckii (Lomnicki, 1894)
- Coelambus ganglbaueri (Lomnicki, 1894)
- Coelambus infernalis (Scudder, 1900)
- Coelambus latefasciatus (Lomnicki, 1894)
- Coelambus miocenus (Wickham, 1912)
- Coelambus niedzwiedzkii (Lomnicki, 1894)
- Coelambus ozokeriticus (Lomnicki, 1894)
- Coelambus picipoides (Lomnicki, 1894)
- Coelhydrus Sharp, 1882
- Colymbetes Clairville, 1806
- Colymbetes Schellenberg, 1806
- Comaldessus Spangler & Barr, 1995
- Copelatus Erichson, 1832
- Copelatus abonnenci Guignot, 1939
- Copelatus acamas Guignot, 1955
- Copelatus adelbert Megna, Atthakor, Manaono, Hendrich & Balke, 2017
- Copelatus advena Sharp, 1882
- Copelatus aemulus Bilardo & Rocchi, 1995
- Copelatus aequatorius Régimbart, 1899
- Copelatus aethiopicus Régimbart, 1906
- Copelatus agaroensis Balke, 2008
- Copelatus agrias Guignot, 1954
- Copelatus alternatus Sharp, 1882
- Copelatus amaroides Guignot, 1952
- Copelatus amatolensis Omer-Cooper, 1965
- Copelatus amazonicus Régimbart, 1889
- Copelatus ambiguus Bertrand & Legros, 1975
- Copelatus amphibius Ranarilalatiana & Bergsten, 2019
- Copelatus anastomosans Guignot, 1952
- Copelatus andobonicus Guignot, 1960
- Copelatus andreinii Régimbart, 1905
- Copelatus andrewesi J.Balfour-Browne, 1939
- Copelatus angolensis Peschet, 1924
- Copelatus angouei Bilardo & Rocchi, 2011
- Copelatus angustus Gschwendtner, 1932
- Copelatus ankaratra Ranarilalatiana & Bergsten, 2019
- Copelatus annobomensis J.Balfour-Browne, 1939
- Copelatus anthracinus Régimbart, 1895
- Copelatus antoniorum Hájek & Brancucci, 2011
- Copelatus aphroditae Balke, 2003
- Copelatus apicinotatus Bilardo & Rocchi, 2018
- Copelatus assalyi Bilardo & Rocchi, 2010
- Copelatus assimilis Régimbart, 1895
- Copelatus ateles Guignot, 1955
- Copelatus atrosulcatus Régimbart, 1906
- Copelatus babyrousa Hájek, Hendrich & Balke, 2018
- Copelatus bacchusi Wewalka, 1981
- Copelatus bacillifer Guignot, 1955
- Copelatus baculiformis Guignot, 1955
- Copelatus badeni Sharp, 1882
- Copelatus bakewelli J.Balfour-Browne, 1939
- Copelatus baoulicus Bilardo & Pederzani, 1978
- Copelatus bapounouensis Bilardo & Rocchi, 2019
- Copelatus baranensis Hájek, Shaverdo, Hendrich & Balke, 2021
- Copelatus barbouri Young, 1942
- Copelatus basifasciatus Régimbart, 1895
- Copelatus basilewskyi Bilardo & Pederzani, 1979
- Copelatus bechynei Guignot, 1953
- Copelatus befasicus Guignot, 1956
- Copelatus bengalensis Guignot, 1955
- Copelatus bertrandi Nilsson, Bilardo & Rocchi, 1996
- Copelatus betampona Ranarilalatiana & Bergsten, 2019
- Copelatus bezdeki Sheth, Ghate & Hájek, 2018
- Copelatus bibulus Guignot, 1948
- Copelatus bicolor (Fabricius, 1792)
- Copelatus biformis Sharp, 1882
- Copelatus bimaculatus (Resende & Vanin, 1991)
- Copelatus binaghii Bilardo & Pederzani, 1978
- Copelatus biolleyi Guignot, 1952
- Copelatus biroi Guignot, 1956
- Copelatus birougouensis Bilardo & Rocchi, 2018
- Copelatus biseriatus J.Balfour-Browne, 1939
- Copelatus biswasi Mukherjee & Sengupta, 1986
- Copelatus blancasi Guignot, 1958
- Copelatus blatchleyi Young, 1953
- Copelatus bolivianus Guignot, 1957
- Copelatus bombycinus Guignot, 1952
- Copelatus bonvouloiri Sharp, 1882
- Copelatus bottegoi Régimbart, 1895
- Copelatus bougainvillensis Hájek, Shaverdo, Hendrich & Balke, 2021
- Copelatus boukali Hendrich & Balke, 1998
- Copelatus brancuccii Rocchi, 1979
- Copelatus brasiliensis Zimmermann, 1921
- Copelatus brendelli Hájek, Hendrich & Balke, 2018
- Copelatus brevicornis (Sharp, 1882)
- Copelatus brevistrigatus Guignot, 1959
- Copelatus brivioi Rocchi, 1976
- Copelatus bromeliarum (Scott, 1912)
- Copelatus brullei Aubé, 1838
- Copelatus brunneus J.Balfour-Browne, 1939
- Copelatus buqueti Aubé, 1838
- Copelatus burgeoni Gschwendtner, 1930
- Copelatus caelatipennis Aubé, 1838
- Copelatus caelatipennis Chevrolat, 1863
- Copelatus caelatus Guignot, 1952
- Copelatus caffer J.Balfour-Browne, 1939
- Copelatus calaquei Bilardo & Rocchi, 2008
- Copelatus camerunensis Guignot, 1941
- Copelatus capensis Sharp, 1882
- Copelatus carayoni Legros, 1949
- Copelatus carinatus Sharp, 1882
- Copelatus celinoides Guignot, 1952
- Copelatus cessaima Caetano, Bená & Vanin, 2013
- Copelatus chevrolati Aubé, 1838
- Copelatus chibcha Guignot, 1952
- Copelatus chinensis Regimbart, 1899
- Copelatus chipiriricus Guignot, 1957
- Copelatus chloroticus Régimbart, 1899
- Copelatus cinnamomeus Régimbart, 1895
- Copelatus clarki Sharp, 1882
- Copelatus collarti Gschwendtner, 1932
- Copelatus compertus Guignot, 1956
- Copelatus concii Bilardo, 1982
- Copelatus concolor Sharp, 1882
- Copelatus concolorans J.Balfour-Browne, 1939
- Copelatus confinis Bilardo & Rocchi, 1999
- Copelatus congo Gschwendtner, 1938
- Copelatus consimilis Bilardo & Rocchi, 2002
- Copelatus consors Sharp, 1882
- Copelatus cooperae Bilardo & Pederzani, 1972
- Copelatus cordovai Megna & Epler, 2012
- Copelatus cordylinoides J.Balfour-Browne, 1938
- Copelatus coxalis Sharp, 1882
- Copelatus crassus Régimbart, 1895
- Copelatus cryptarchoides Régimbart, 1899
- Copelatus cubaensis Schaeffer, 1908
- Copelatus curtistriatus Bilardo & Rocchi, 1995
- Copelatus curvispinis Bilardo & Rocchi, 2008
- Copelatus cynthiae Bilardo & Rocchi, 2008
- Copelatus daemeli Sharp, 1882
- Copelatus danyi Megna & Epler, 2012
- Copelatus darlingtoni Young, 1942
- Copelatus davidi Wewalka, 2017
- Copelatus debilis Sharp, 1882
- Copelatus deccanensis Sheth, Ghate & Hájek, 2018
- Copelatus decellei Bilardo, 1982
- Copelatus decemsulcatus Régimbart, 1895
- Copelatus deceptor Bilardo & Rocchi, 1995
- Copelatus dentatipenis Jiang, Hajek & Jia, 2022
- Copelatus depressus Sharp, 1882
- Copelatus descarpentriesi Bertrand & Legros, 1975
- Copelatus diffisus Guignot, 1939
- Copelatus dimorphus Sharp, 1882
- Copelatus distinctus Aubé, 1838
- Copelatus distinguendus Régimbart, 1903
- Copelatus diversistriatus Jiang, Hajek & Jia, 2022
- Copelatus djiboutensis Wewalka & Jäch, 2017
- Copelatus dolosus Guignot, 1956
- Copelatus doriae Sharp, 1882
- Copelatus doudouensis Bilardo & Rocchi, 2015
- Copelatus duodecimstriatus Aubé, 1838
- Copelatus duponti Aubé, 1838
- Copelatus efoutensis Bilardo & Rocchi, 1995
- Copelatus ejactus Omer-Cooper, 1965
- Copelatus ellai Bilardo & Rocchi, 1995
- Copelatus elutus Guignot, 1958
- Copelatus enganensis Guignot, 1940
- Copelatus epactus Guignot, 1948
- Copelatus erichsonii Guérin-Méneville, 1847
- Copelatus espinhasso Hájek, Alarie, Benetti, Hamada, Springer, Hendrich, Villastrigo, Torres, Basantes & Balke, 2024[7]
- Copelatus esteriensis Bilardo & Pederzani, 1978
- Copelatus eucritus Guignot, 1952
- Copelatus evanidus Bilardo & Rocchi, 1995
- Copelatus exaratus Sharp, 1882
- Copelatus externus Kirsch, 1873
- Copelatus falciformis Pederzani & Schizzerotto, 2017
- Copelatus fasciatus Bilardo & Rocchi, 1995
- Copelatus fastidiosus Guignot, 1959
- Copelatus feae Régimbart, 1888
- Copelatus felicis Hajek, Jiang & Jia, 2022
- Copelatus fernandensis Guignot, 1952
- Copelatus ferruginicollis Régimbart, 1895
- Copelatus ferus Guignot, 1953
- Copelatus festae Régimbart, 1899
- Copelatus fidschiensis Zimmermann, 1928
- Copelatus fijiensis Guignot, 1955
- Copelatus filiformis Sharp, 1882
- Copelatus fizpaci Bilardo & Rocchi, 2013
- Copelatus flavicans Guignot, 1952
- Copelatus flavidus Régimbart, 1895
- Copelatus florae Hájek, Alarie, Benetti, Hamada, Springer, Hendrich, Villastrigo, Torres, Basantes & Balke, 2024[7]
- Copelatus fluviaticus Guignot, 1957
- Copelatus fontanus J.Balfour-Browne, 1950
- Copelatus fossilis Říha, 1974
- Copelatus fractistriatus Bilardo & Rocchi, 1995
- Copelatus freudei Guignot, 1955
- Copelatus fryi J.Balfour-Browne, 1939
- Copelatus fulviceps J.Balfour-Browne, 1938
- Copelatus fuscipennis Sharp, 1882
- Copelatus fuscomaculatus Guignot, 1952
- Copelatus gabonicus Bilardo & Pederzani, 1978
- Copelatus galapagoensis G.R.Waterhouse, 1845
- Copelatus garambanus Guignot, 1955
- Copelatus gardineri Scott, 1912
- Copelatus geayi Régimbart, 1904
- Copelatus geniculatus Sharp, 1882
- Copelatus gentilis Sharp, 1882
- Copelatus gestroi (Sharp, 1882)
- Copelatus gigas Bilardo & Rocchi, 2019
- Copelatus glyphicus (Say, 1823)
- Copelatus griffinii Régimbart, 1899
- Copelatus gschwendtneri Guignot, 1939
- Copelatus guadelupensis Legros, 1948
- Copelatus guerini Aubé, 1838
- Copelatus guineensis Guignot, 1955
- Copelatus haemorrhoidalis Régimbart, 1883
- Copelatus hararensis Guignot, 1952
- Copelatus hardenbergi J.Balfour-Browne, 1950
- Copelatus hebeter Guignot, 1953
- Copelatus heterogynus Régimbart, 1899
- Copelatus hydroporoides (Murray, 1859)
- Copelatus ibrahimi Zalat, Saleh, Angus & Kaschef, 2000
- Copelatus iguelaensis Bilardo & Rocchi, 2002
- Copelatus ilybioides Régimbart, 1895
- Copelatus imitator Bilardo & Rocchi, 2002
- Copelatus inaequalis Sharp, 1882
- Copelatus incognitus Sharp, 1882
- Copelatus indicus Sharp, 1882
- Copelatus inopinatus Bilardo & Rocchi, 1995
- Copelatus inornatus Sharp, 1882
- Copelatus insidiosus Bilardo & Rocchi, 1995
- Copelatus insolitus Chevrolat, 1863
- Copelatus instabilis Régimbart, 1897
- Copelatus instriatus Zimmermann, 1921
- Copelatus insuetus Guignot, 1941
- Copelatus insulanus Guignot, 1939
- Copelatus integer Sharp, 1882
- Copelatus internus Régimbart, 1904
- Copelatus interstriatus J.Balfour-Browne, 1939
- Copelatus inuber Guignot, 1952
- Copelatus ipiformis Régimbart, 1895
- Copelatus irinus Régimbart, 1899
- Copelatus irregularis W.J.MacLeay, 1871
- Copelatus ischius Guignot, 1956
- Copelatus jactator Guignot, 1955
- Copelatus jaechi Bilardo & Rocchi, 2016
- Copelatus jamaicensis Young, 1942
- Copelatus japonicus Sharp, 1884
- Copelatus jarrigei Legros, 1954
- Copelatus javanus Régimbart, 1883
- Copelatus jocosus Guignot, 1955
- Copelatus johannis Guignot, 1954
- Copelatus kalaharii Gschwendtner, 1935
- Copelatus kammuriensis Tamu & Tsukamoto, 1955
- Copelatus kaszabi Guignot, 1956
- Copelatus kely Ranarilalatiana & Bergsten, 2019
- Copelatus kietensis Hájek, Shaverdo, Hendrich & Balke, 2021
- Copelatus kilimandjarensis Bilardo & Pederzani, 1972
- Copelatus kindianus Guignot, 1954
- Copelatus kongouensis Bilardo & Rocchi, 1999
- Copelatus koreanus Mori, 1932
- Copelatus laccophilinus Sharp, 1882
- Copelatus laeticulus Sharp, 1882
- Copelatus laevipennis Hájek, Shaverdo, Hendrich & Balke, 2021
- Copelatus lamottei Legros, 1954
- Copelatus lanzai Bilardo & Rocchi, 1995
- Copelatus laraensis Bilardo & Rocchi, 1995
- Copelatus lasckonyi Bilardo & Rocchi, 1995
- Copelatus latens Guignot, 1952
- Copelatus laticollis Régimbart, 1899
- Copelatus latifasciatus Bilardo & Rocchi, 1999
- Copelatus latipes Sharp, 1882
- Copelatus latus J.Balfour-Browne, 1950
- Copelatus leonardii Bilardo & Rocchi, 1999
- Copelatus leonensis Legros, 1954
- Copelatus lepersonneae Gschwendtner, 1943
- Copelatus lignosus Guignot, 1955
- Copelatus lineatipennis Guignot, 1955
- Copelatus lineatus (Guérin-Méneville, 1838)
- Copelatus longicornis Sharp, 1882
- Copelatus lootensi Guignot, 1955
- Copelatus louayensis Bilardo & Rocchi, 2004
- Copelatus luctuosus Guignot, 1939
- Copelatus luridescens Régimbart, 1889
- Copelatus luteocinctus Guignot, 1955
- Copelatus luteomaculatus Guignot, 1956
- Copelatus luzonicus Guignot, 1952
- Copelatus macellus Guignot, 1950
- Copelatus madoni Guignot, 1952
- Copelatus mahajanga Pederzani & Hájek, 2005
- Copelatus mahleri Holmen & Vazirani, 1990
- Copelatus makokouensis Bilardo & Rocchi, 1995
- Copelatus malaisei Guignot, 1954
- Copelatus malavergnorum Manuel & Ramahandrison, 2022
- Copelatus mancus Sharp, 1887
- Copelatus marginatus Sharp, 1882
- Copelatus marginipennis (Laporte, 1835)
- Copelatus martinbaehri Hendrich, Shaverdo, Hájek & Balke, 2019
- Copelatus martini Manuel, Deler-Hernández, Megna & Hájek, 2018
- Copelatus masculinus Régimbart, 1899
- Copelatus massaicus Guignot, 1941
- Copelatus mathani Guignot, 1952
- Copelatus maushomi Sheth, Ghate & Hájek, 2018
- Copelatus mbokoensis Bilardo & Rocchi, 2006
- Copelatus melanogrammus Régimbart, 1883
- Copelatus mendax Bilardo & Rocchi, 2011
- Copelatus miaowangi Hajek, Jiang & Jia, 2022
- Copelatus minimus Bilardo & Rocchi, 1995
- Copelatus minor Bilardo & Pederzani, 1978
- Copelatus minutissimus J.Balfour-Browne, 1939
- Copelatus mocquerysi Régimbart, 1895
- Copelatus mohelicus Guignot, 1959
- Copelatus monticola Guignot, 1951
- Copelatus montivagus Young, 1942
- Copelatus mopanshanensis Jiang, Zhao & Hajek, 2022
- Copelatus mulangensis Bameul, 2003
- Copelatus mundus Sharp, 1882
- Copelatus mutabilis Bilardo & Rocchi, 1999
- Copelatus mvoungensis Bilardo & Rocchi, 2004
- Copelatus mysorensis Vazirani, 1970
- Copelatus nakamurai Guéorguiev, 1970
- Copelatus nangaensis Guignot, 1952
- Copelatus nanlingensis Hajek, Jiang & Jia, 2022
- Copelatus nauclerus Guignot, 1939
- Copelatus neavei J.Balfour-Browne, 1950
- Copelatus neelumae Vazirani, 1973
- Copelatus neogestroi Balke, 2008
- Copelatus neoguineensis Zimmermann, 1919
- Copelatus nguemai Bilardo & Rocchi, 2011
- Copelatus nigricans Sharp, 1882
- Copelatus nigricollis Paulino de Oliveira, 1880
- Copelatus nigrolineatus Sharp, 1882
- Copelatus nigropennis Zimmermann, 1927
- Copelatus nigrostriatus Régimbart, 1895
- Copelatus nilssoni Wewalka, 1982
- Copelatus nimbaensis Legros, 1958
- Copelatus nitens Bilardo & Rocchi, 1999
- Copelatus nitidus Sharp, 1882
- Copelatus nodieri Régimbart, 1895
- Copelatus normalis Erichson, 1847
- Copelatus notius Omer-Cooper, 1965
- Copelatus nzei Bilardo & Rocchi, 1999
- Copelatus oblitus Sharp, 1882
- Copelatus obscurus Sharp, 1882
- Copelatus occultus Bilardo & Rocchi, 1995
- Copelatus ogasawarensis Kamiya, 1932
- Copelatus onorei Pederzani & Rocchi, 1982
- Copelatus ornatipennis Zimmermann, 1926
- Copelatus owas Régimbart, 1895
- Copelatus pachys Guignot, 1955
- Copelatus pallidus Régimbart, 1895
- Copelatus paludorum Guignot, 1952
- Copelatus pandanorum Scott, 1912
- Copelatus panguana Hájek, Alarie, Benetti, Hamada, Springer, Hendrich, Villastrigo, Torres, Basantes & Balke, 2024[7]
- Copelatus pantosi Guignot, 1958
- Copelatus papuensis J.Balfour-Browne, 1939
- Copelatus parabaptus Guignot, 1955
- Copelatus parallelipipedus Régimbart, 1895
- Copelatus parallelus Zimmermann, 1920
- Copelatus pardii Rocchi, 1990
- Copelatus parisii Guignot, 1959
- Copelatus parumstriatus Gschwendtner, 1934
- Copelatus paryphes Guignot, 1955
- Copelatus pederzanii Bilardo & Rocchi, 1995
- Copelatus pereirai Guignot, 1955
- Copelatus peridinus Guignot, 1955
- Copelatus pinnifer Guignot, 1952
- Copelatus piriensis Omer-Cooper, 1965
- Copelatus platynotus Régimbart, 1895
- Copelatus polystrigus Sharp, 1882
- Copelatus ponomarenkoi Říha, 1974
- Copelatus portior Guignot, 1956
- Copelatus posticatus (Fabricius, 1801)
- Copelatus poungai Bilardo & Rocchi, 2006
- Copelatus predaveterus Miller, 2003
- Copelatus prolixus Sharp, 1882
- Copelatus prolongatus Sharp, 1882
- Copelatus propino Guignot, 1960
- Copelatus propinquus Régimbart, 1895
- Copelatus proximus Sharp, 1882
- Copelatus pseudostriatus Ranarilalatiana & Bergsten, 2019
- Copelatus pulchellus (Klug, 1834)
- Copelatus pulicarius Régimbart, 1895
- Copelatus pumilus Régimbart, 1895
- Copelatus punctatus Bilardo & Rocchi, 1995
- Copelatus punctulatus Aubé, 1838
- Copelatus puzhelongi Jiang, Hajek & Jia, 2022
- Copelatus quadrisignatus Régimbart, 1877
- Copelatus quadristriatus Pederzani & Schizzerotto, 2017
- Copelatus racenisi Guignot, 1951
- Copelatus ragazzii Régimbart, 1887
- Copelatus regimbarti Branden, 1885
- Copelatus restrictus Sharp, 1882
- Copelatus rimosus Guignot, 1952
- Copelatus rivalis Guignot, 1952
- Copelatus rocchii Bilardo, 1982
- Copelatus romani Zimmermann, 1924
- Copelatus rosulae Hajek, Jiang & Jia, 2022
- Copelatus royi Legros, 1958
- Copelatus rubiginosus Guignot, 1954
- Copelatus ruficapillus Régimbart, 1895
- Copelatus ruteri Legros, 1954
- Copelatus saegeri Guignot, 1955
- Copelatus safiotra Ranarilalatiana & Bergsten, 2019
- Copelatus sahlbergi J.Balfour-Browne, 1939
- Copelatus sallaei Sharp, 1882
- Copelatus sanfilippoi Bilardo & Pederzani, 1972
- Copelatus saverii Pederzani & Schizzerotto, 2017
- Copelatus scalptus Guignot, 1954
- Copelatus scaphites Guignot, 1955
- Copelatus schaefferi Young, 1942
- Copelatus schereri Wewalka, 1981
- Copelatus schuhi Hendrich & Balke, 1998
- Copelatus scutatus Guignot, 1952
- Copelatus scytalotus Guignot, 1956
- Copelatus senegalensis Lagar & Hernando, 1991
- Copelatus sexstriatus Sharp, 1882
- Copelatus sharpi Branden, 1885
- Copelatus sibelaemontis Hájek, Hendrich, Hawlitschek & Balke, 2010
- Copelatus silvestrii Régimbart, 1903
- Copelatus simoni Régimbart, 1889
- Copelatus singularis Bilardo & Rocchi, 1995
- Copelatus sociennus J.Balfour-Browne, 1952
- Copelatus solitarius Sharp, 1882
- Copelatus sordidipennis Régimbart, 1895
- Copelatus speciosus Régimbart, 1892
- Copelatus spoliatus Guignot, 1955
- Copelatus stavropolitanus Říha, 1974
- Copelatus stillicidicola Bilardo & Rocchi, 1995
- Copelatus striatellus Boheman, 1848
- Copelatus striaticollis Lucas, 1857
- Copelatus striatopterus Aubé, 1838
- Copelatus striatulus Aubé, 1838
- Copelatus strigipennis (Laporte, 1835)
- Copelatus strigosulus (Fairmaire, 1878)
- Copelatus strinatii Guignot, 1958
- Copelatus striolatus Peschet, 1917
- Copelatus stygis Guignot, 1958
- Copelatus subdeficiens Régimbart, 1902
- Copelatus subsimilis Guignot, 1958
- Copelatus substriatus Kirsch, 1873
- Copelatus subterraneus Guéorguiev, 1978
- Copelatus sudrei Bameul, 2003
- Copelatus sulcatus Sharp, 1882
- Copelatus sulcipennis (Laporte, 1835)
- Copelatus sumbawensis Régimbart, 1899
- Copelatus suppar Guignot, 1956
- Copelatus supplementaris Régimbart, 1895
- Copelatus sylvaticoides Bilardo & Rocchi, 2013
- Copelatus sylvaticus Guignot, 1952
- Copelatus takakurai Satô, 1985
- Copelatus tanaus Guignot, 1953
- Copelatus taprobanicus Wewalka & Vazirani, 1985
- Copelatus tenebrosis Regimbart, 1880
- Copelatus tenebrosus Régimbart, 1880
- Copelatus tengchongensis Hajek, Jiang & Jia, 2022
- Copelatus teranishii Kamiya, 1938
- Copelatus terminalis Sharp, 1882
- Copelatus ternatensis Régimbart, 1899
- Copelatus terzanii Bilardo & Saverio, 2016
- Copelatus thiriouxi Peschet, 1917
- Copelatus thrasys Guignot, 1952
- Copelatus tibialis Sharp, 1882
- Copelatus tinctor Guignot, 1954
- Copelatus togoensis Régimbart, 1895
- Copelatus tomokunii Satô, 1985
- Copelatus tondangoyei Bilardo & Rocchi, 2013
- Copelatus tostus J.Balfour-Browne, 1950
- Copelatus trifilis Guignot, 1958
- Copelatus triglyphus Guignot, 1955
- Copelatus trilobatus Régimbart, 1895
- Copelatus tschaga Bilardo & Pederzani, 1972
- Copelatus tucuchiensis J.Balfour-Browne, 1939
- Copelatus tulagicus Guignot, 1942
- Copelatus uludanuensis Hendrich & Balke, 1995
- Copelatus undecimstriatus Aubé, 1838
- Copelatus urceolus Hájek, Shaverdo, Hendrich & Balke, 2021
- Copelatus usagarensis Zimmermann, 1926
- Copelatus vagatus Guignot, 1952
- Copelatus vagestriatus Zimmermann, 1919
- Copelatus vagus Bilardo & Rocchi, 1995
- Copelatus validus Sharp, 1882
- Copelatus vancli Hájek, Hendrich & Balke, 2018
- Copelatus vanninii Bilardo & Rocchi, 1999
- Copelatus variegatus Régimbart, 1895
- Copelatus variistriatus Hájek, Shaverdo, Hendrich & Balke, 2021
- Copelatus venustus Bilardo & Rocchi, 1995
- Copelatus vigintistriatus Fairmaire, 1869
- Copelatus vigintisulcatus Régimbart, 1895
- Copelatus villiersi Guignot, 1950
- Copelatus virungaensis Bilardo, 1982
- Copelatus vitraci Legros, 1948
- Copelatus vivax Guignot, 1953
- Copelatus vokoka Ranarilalatiana & Bergsten, 2019
- Copelatus wallacei J.Balfour-Browne, 1939
- Copelatus waltoni J.Balfour-Browne, 1950
- Copelatus wayanadensis Manivannan & Madani, 2011
- Copelatus wewalkai Holmen & Vazirani, 1990
- Copelatus weymarni J.Balfour-Browne, 1947
- Copelatus xanthocephalus Régimbart, 1899
- Copelatus xiniudong Jiang, Hajek & Jia, 2022
- Copelatus yacumensis Guignot, 1957
- Copelatus yaguarete Scheers, 2018
- Copelatus yombensis Bilardo & Rocchi, 2015
- Copelatus zadiensis Bilardo & Rocchi, 1995
- Copelatus zanabato Manuel & Ramahandrison, 2022
- Copelatus zanatanensis Ranarilalatiana & Bergsten, 2019
- Copelatus zimmermanni Gschwendtner, 1934
- Copelatus Sheth, Ghate & Hájek, 2018
- Coptotomus Say, 1830
- Cretodytes Ponomarenko, 1977
- Crinodessus K.B.Miller, 1997
- 大龍蝨屬 Cybister Curtis, 1827，真龙虱属
- Darwinhydrus Sharp, 1882
- Deronectes Sharp, 1882
- Derovatellus Sharp, 1882
- Desmopachria Babington, 1841
- Deuteronectes Guignot, 1945
- Deuteronectus
- Dimitshydrus Uéno, 1996
- Dyschirioides
- Dyticus Geoffroy, 1762
- Dytiscus Linnaeus, 1758
- Electruphilus Balker, Toledo & Hendrich, 2019
- Ereboporus K.B.Miller, Gibson & Alarie, 2009
- Eretes Laporte, 1833
- Etruscodytes Mazza, Cianferoni & Rocchi, 2013
- Exocelina Broun, 1886
- Fontidessus K.B.Miller & Spangler, 2008
- Geodessus Brancucci, 1979
- Gibbidessus Watts, 1978
- Glareadessus Wewalka & Biström, 1998
- Graphoderus Dejean, 1833
- Graptodytes Seidlitz, 1887
- Haideoporus Young & Longley, 1976
- Hamadiana Benetti, Short & Michat, 2019
- Hemibidessus Zimmermann, 1921
- Heroceras Guignot, 1950
- Heterhydrus Fairmaire, 1869
- Heterosternuta Strand, 1935
- Hintonia Young, 1980
- Hoperius Fall, 1927
- Hornectes Fery & Ribera, 2018
- Hovahydrus Biström, 1982
- Huxelhydrus Sharp, 1882
- Hydaticus Leach, 1817
- Hyderodes Hope, 1838
- Hydrocolus Roughley & Larson, 2000
- Hydrodessus J.Balfour-Browne, 1953
- Hydrodytes K.B.Miller, 2001
- 短褶龙虱属 Hydroglyphus Motschulsky, 1853
- Hydronebrius Jakovlev, 1897
- Hydropeplus Sharp, 1882
- Hydroporus Clairville, 1806
- Hydroporus Schellenberg, 1806
- Hydrotrupes Sharp, 1882
- Hydrovatus Motschulsky, 1853
- 水龙虱属 Hygrotus Stephens, 1828
- Hyphoporus Sharp, 1882
- Hyphovatus Wewalka & Biström, 1994
- 异爪龙虱属 Hyphydrus Illiger, 1802
- Hypodessus Guignot, 1939
- Iberonectes Fery & Ribera, 2018
- Iberoporus Castro & Delgado, 2001
- Ilybiosoma Crotch, 1873
- Ilybius Erichson, 1832
- Incomptodessus K.B.Miller & García, 2011
- Isonotus Houlbert, 1934
- Japanolaccophilus Satô, 1972
- Kakadudessus Hendrich & Balke, 2009
- Kakadudessus tomweiri Hendrich & Balke, 2009
- Kuschelydrus Ordish, 1976
- Laccodytes Régimbart, 1895
- Laccomimus Toledo & Michat, 2015
- Lacconectus Motschulsky, 1855
- 粒龙虱属 Laccophilus Leach, 1815
- Laccoporus J.Balfour-Browne, 1939
- Laccoporus nigritulus (Gschwendtner, 1936)
- Laccornellus Roughley & Wolfe, 1987
- Laccornis Gozis, 1914
- Laccosternus Brancucci, 1983
- Lancetes Sharp, 1882
- Laodytes Queney, Lemaire & Ferrand, 2020
- Larsonectes Fery & Ribera, 2018
- Leconectes Fery & Ribera, 2018
- Leiodytes Guignot, 1936
- Leuronectes Sharp, 1882
- Liadyxianus Prokin, Petrov, Wang & Ponomarenko, 2013
- Limbodessus Guignot, 1939
- Liodessus Guignot, 1939
- Lioporeus Guignot, 1950
- Liopterus Dejean, 1833
- Madaglymbus Shaverdo & Balke, 2008
- Matus Aubé, 1836
- Megadytes Sharp, 1882
- Megaporus Brinck, 1943
- Meladema Laporte, 1835
- Melanodytes Seidlitz, 1887
- Meridiorhantus Balke, Hájek & Hendrich, 2017
- Mesodytes Prokin, Petrov, Wang & Ponomarenko, 2013
- Metaporus Guignot, 1945
- Methles Sharp, 1882
- Microdessus Young, 1967
- Microdytes J.Balfour-Browne, 1946
- Microhydrodytes K.B.Miller, 2002
- Miodytiscus Wickham, 1911
- Morimotoa Uéno, 1957
- Mystonectes Fery & Ribera, 2018
- Napodytes Steiner, 1981
- Nartus Zaitzev, 1907
- 孔龙虱属 Nebrioporus Régimbart, 1906
- Necterosoma Sharp, 1882
- Necterosoma W.J.MacLeay, 1871
- Nectoboreus Fery & Ribera, 2018
- Nectomimus Fery & Ribera, 2018
- Nectoporus Guignot, 1950
- Neobidessodes Hendrich & Balke, 2009
- Neobidessus Young, 1967
- Neoclypeodytes Young, 1967
- Neonectes J.Balfour-Browne, 1944
- Neonectes Zimmermann, 1932
- Neoporus Guignot, 1931
- Neoscutopterus J.Balfour-Browne, 1943
- 彩裳粒龙虱属 Neptosternus Sharp, 1882
- Notaticus Zimmermann, 1928
- Novadessus K.B.Miller, 2016
- Onychohydrus Schaum & White, 1847
- Oreodytes Seidlitz, 1887
- Pachydrus Sharp, 1882
- Pachynectes Régimbart, 1903
- Palaeodytes Ponomarenko, 1987
- Papuadessus Balke, 2001
- Paroster Sharp, 1882
- Peschetius Guignot, 1942
- Petrodessus K.B.Miller, 2012
- Philaccolilus Guignot, 1937
- Philaccolus Guignot, 1937
- Philodytes J.Balfour-Browne, 1939
- Phreatodessus Ordish, 1976
- Platambus Thomson, 1859
- Platydytes Biström, 1988
- Platynectes Gueorguiev, 1972
- Platynectes Régimbart, 1879
- Porhydrus Guignot, 1945
- Primospes Sharp, 1882
- Procoelambus Théobald, 1937
- Prodaticus Sharp, 1882
- 缝龙虱属 Pseuduvarus Biström, 1988
- Psychopomporus Jean, Telles & K.B.Miller, 2012
- Pteroporus Guignot, 1933
- Queda Sharp, 1882
- Regimbartina Chatanay, 1911
- Rhantaticus Sharp, 1880
- Rhantus Dejean, 1833
- Rhithrodytes Bameul, 1989
- Rompindessus Balke, Bergsten & Hendrich, 2017
- Sandracottus Sharp, 1882
- Sanfilippodytes Franciscolo, 1979
- Scarodytes Gozis, 1914
- Schistomerus Palmer, 1957
- Sekaliporus Watts, 1997
- Sharphydrus Omer-Cooper, 1958
- Siamoporus Spangler, 1996
- Siettitia Abeille de Perrin, 1904
- Sinodytes Spangler, 1996
- Spanglerodessus K.B.Miller & García, 2011
- Spencerhydrus Sharp, 1882
- Sternhydrus Brinck, 1945
- Sternopriscus Sharp, 1880
- Stictonectes Brinck, 1943
- Stictotarsus Zimmermann, 1919
- Stygoporus Larson & LaBonte, 1994
- Suphisellus Zimmermann, 1921
- Tassilodytes Fery & Bouzid, 2016
- Tepuidessus Spangler, 1981
- Thermonectus Dejean, 1833
- Thermonectus alfredi Griffini, 1898
- Thermonectus basillaris (Harris, 1829)
- Thermonectus batesi Sharp, 1882
- Thermonectus circumscriptus (Latreille, 1809)
- Thermonectus cuneatus Sharp, 1882
- Thermonectus depictus Sharp, 1882
- Thermonectus duponti (Aubé, 1838)
- Thermonectus intermedius Crotch, 1873
- Thermonectus laporti (Aubé, 1838)
- Thermonectus leprieuri J.Balfour-Browne, 1944
- Thermonectus margineguttatus (Aubé, 1838)
- Thermonectus marmoratus (Gray, 1831)
- Thermonectus nigrofasciatus (Aubé, 1838)
- Thermonectus nobilis Zimmermann, 1924
- Thermonectus sibleyi Goodhue-McWilliams, 1981
- Thermonectus simulator Sharp, 1882
- Thermonectus succinctus (Aubé, 1838)
- Thermonectus tremouillesi Michat & Torres, 2016[8]
- Thermonectus variegatus (Dejean, 1836)
- Thermonectus variegatus (Laporte de Castelnau, 1835)
- Thermonectus zimmermani Goodhue-McWilliams, 1981
- Tikoloshanes Omer-Cooper, 1956
- Tiporus Watts, 1985
- Trichonectes Guignot, 1941
- Trogloguignotus Sanfilippo, 1958
- Tyndallhydrus Sharp, 1882
- Typhlodessus Brancucci, 1985
- Uvarus Guignot, 1939
- Vatellus Aubé, 1837
- Yola Gozis, 1886
- Yolina Guignot, 1936
- Zaitzevhydrus Fery & Ribera, 2018
- Zimpherus K.B.Miller & Wheeler, 2015
參考文獻
- 联邦科学与工业研究组织 (CSIRO) (2004): Water for a Healthy Country - Family Dytiscidae （页面存档备份，存于互联网档案馆）. Version of 2004-JUL-02. Retrieved 2008-AUG-04
- De Foliart, Gene R. (2002): Chapter 26 - Eastern Asia: China, Japan, and other countries. In: The Human Use of Insects as a Food Resource: A Bibliographic Account in Progress.
- Jäch, Manfred A. (2003): Fried water beetles Cantonese style. American Entomologist 49(1): 34-37. PDF fulltext
- Larson, D.J., Alarie, Y., and Roughley, R.E. (2000): Predaceous Diving Beetles (Coleoptera: Dytiscidae) of the Nearctic Region, with emphasis on the fauna of Canada and Alaska. NRC Research Press, Ottawa. ISBN 978-0-660-17967-4.
